組3. 翠雀花組 Sect. Delphinastrum

Sect. Delphinastrum DC. Syst. 1： 351. 1818; et Prodr. 1： 53. 1824; W. T. Wang in Acta Bot. Sin. 10（1）： 155. 1962; Tamura in Sei. Rep. Osaka Univ. 15： 33. 1966; et in Hiepko， Nat. Pflanzenfam.， Zwei. Aufl.， 17aIV： 299. 1995; W. T. Wang & M. J. Warnock in Fl. China 6： 246. 2001. Lectotype： Delphinium grandiflorum L. （Wang， 1962）.

Sect. Schizopetala Huth in Bull. Herb. Boiss. 1： 333. 1893.

Sect.Diedropetala Huth in Bot. Jahrb. 20： 420. 1895. — Gen. Diedropetala （Huth） Galushko in Novost. Syst. Pl. Vasc. 15： 251. 1976.

Sect. Kalobopetala Huth in l. c. 458

Sect. Homochroe Bruhl in Ann. Bot. Gard. Calc. 5： 91. 1896.

Sect. Isocheilae Bruhl in l. c.， p. p.

不具塊根。葉3淺裂至3全裂。花序總狀或傘房狀，或花單朵頂生。萼片脫落，藍(lán)色或紫色，稀白色，多少展開，稀上萼片船形;距與萼片近等長或比萼片長，稀比萼片短?；ò昱c萼片同色，稀淡黃色或黃白色。退化雄蕊瓣片，也與萼片同色，稀淡黃色或黃白色，2淺裂或2半裂，稀不分裂，腹面基部之上被黃色或白色髯毛。心皮3，稀5。種子四面體形，沿縱棱具狹翅或無翅，或狹倒卵球形，具密集橫向排列的小鱗片。

約190種，隸屬于6亞組，廣布于亞洲、歐洲東北部、非洲和北美洲。我國有6亞組約132種，多數(shù)種分布于西南部的四川、云南、西藏、青海等?。▍^(qū)），西部、北部和東北部諸省各有4種至1種。

分種和變種檢索表

1. 花瓣和退化雄蕊淡黃色或淡黃白色;萼片藍(lán)色或紫色。

2. 葉片3全裂;花單朵頂生;萼片比距長;花瓣淡黃白色;退化雄蕊瓣片淡黃白色，不分裂;心皮5

159. 湟中翠雀花Delphinium huangzhongense

2. 葉片3深裂;花組成總狀花序;退化雄蕊瓣片2淺裂;心皮3或1。

3. 萼片與距近等長;花瓣和退化雄蕊淡黃色;心皮3。

4. 花較小，組成復(fù)總狀花序;萼片長6～7 mm，頂端以直角向內(nèi)彎曲;心皮無毛

160. 征鎰翠雀花 D. zhengyianum

4. 花較大，組成簡單總狀花序;萼片長14 mm，不彎曲;子房被短柔毛

161. 文采新翠雀花 D. neowentsaii

3. 萼片比距短;花瓣和退化雄蕊黃白色;心皮1，無毛

189. 單蕊翠雀花 D. monogynum

1. 花瓣和退化雄蕊與萼片同色，即呈藍(lán)色或紫色，稀白色。

5. 基生葉多不存在，或存在，則在花期枯萎。

6. 種子具3縱棱，沿棱具狹翅。

7. 心皮4～6

8. 葉片3全裂;心皮5

98. 密葉翠雀花 D. kingianum

9. 莖有毛;葉全裂片頂端急尖。

10. 莖密被白色短柔毛和黃色短腺毛。

11. 子房密被短柔毛

98a. 模式變種 var. kingianum

11. 子房只在腹縫線上有毛，其他部分無毛

98b. 光果密葉翠雀花 var. leiocarpum

10. 莖密被白色短柔毛，具少數(shù)黃色腺毛

98c. 少腺密葉翠雀花 var. eglandulosum

9. 莖無毛;葉全裂片頂端長漸尖

98d. 尖裂密葉翠雀花 var. acuminatissimum

8. 葉片3深裂;心皮4～6

99. 鄉(xiāng)城翠雀花 D. humilius

7. 心皮3。

12. 葉片3全裂;萼片比距長

100. 尾裂翠雀花 D. caudatolobum

12. 葉片3深裂;萼片與距近等長，或比距短。

13. 總狀花序軸和花梗被白色短柔毛和黃色短腺毛

103. 拉薩翠雀花 D. gyalanum

13. 總狀花序軸和花梗無黃色腺毛。

14. 總狀花序軸和莖無毛。

15. 植株高達(dá)140 cm;萼片長14～15 mm;退化雄蕊瓣片2淺裂

106. 唇花翠雀花 D. cheilanthum

16. 花梗被白色反曲短柔毛

106a. 模式變種 var. cheilanthum

16. 花梗被黃色伸展短毛

106b. 展毛唇花翠雀花 var. pubescens

15. 植株高達(dá)20～30 cm;萼片長15～20 mm。

17. 葉側(cè)生1回裂片2半裂;萼片長15～20 mm，距長18～24 mm;退化雄蕊瓣片近圓形， 2淺裂

104. 寬片翠雀花 D. latilimbum

17. 葉側(cè)生1回裂片2深裂超過葉片中部;萼片長15～18 mm，距長20 mm;退化雄蕊瓣片寬橢圓形，頂端微凹

107. 毓泉翠雀花 D. yuchuanii

14. 花梗和莖有毛，稀莖近無毛 （D. hsinganense）。

18. 萼片兩面均有毛，與距等長或稍長

101. 粗距翠雀花 D. pachycentrum

19. 萼片長圓形

101a. 模式變種var. pachycentrum

19. 萼片披針形

101b. 狹萼粗距翠雀花var. lancisepalum

18. 萼片內(nèi)面無毛。

20. 花序復(fù)總狀;小苞片貼伏于花上，狹披針形，長6～11 mm;萼片長13 mm;退化雄蕊瓣片倒卵狀長圓形，有白色髯毛

102. 擬粗距翠雀花D. pachycentroides

20. 花序簡單總狀;小苞片與花離開，鉆形，長3～5 mm;萼片長14～17 mm;退化雄蕊瓣片圓卵形，有黃色髯毛

105. 興安翠雀花 D. hsinganense

6. 種子具密集橫向排列的小鱗片或橫波狀條紋。

21. 葉片寬心形，不分裂或不明顯3淺裂

158. 心葉翠雀花 D. calthifolium

21. 葉片五角形或腎形，3淺裂，3深裂或3全裂。

22. 葉片3淺裂或3深裂。

23. 葉片3淺裂，分裂不超過葉片中部。

24. 葉片3淺裂不達(dá)葉片中部，淺裂片互相不覆壓;總狀花序長10 cm;花梗被白色柔毛和黃色腺毛

135. 葡萄葉翠雀花 D. sinovitifolium

24. 葉片3淺裂達(dá)葉片中部，淺裂片互相覆壓;傘房花序長6 cm;花梗無毛

134. 茂縣翠雀花 D. maoxianense

23. 葉片3深裂超過中部，達(dá)近基部。

25. 花序傘房狀。

26. 莖無毛。

27. 萼片背面頂端之下具角狀突起

133. 叉角翠雀花 D. furcatocornutum

27. 萼片無角狀突起。

28. 萼距螺旋狀彎曲

137. 川黔翠雀花 D. bonvalotii

28. 萼距稍向下彎曲

130. 毛梗翠雀花 D. eriostylum

26. 莖有毛。

29. 莖被短柔毛;葉片寬達(dá)6 cm

131. 平武翠雀花 D. pingwuense

29. 莖被糙硬毛;葉片寬達(dá)12 cm。

30. 花梗有黃色腺毛

130. 毛梗翠雀花D. eriostylum

31. 花梗只被黃色腺毛;子房疏被柔毛

130a. 模式變種var. eriostylum

31. 花梗被黃色短腺毛和白色短柔毛;心皮無毛

130b. 糙葉毛梗翠雀花 var. hispidum

30. 花梗無黃色腺毛

132. 疏傘翠雀花D. laxicymosum

32. 花梗無毛;萼距長23～27 mm

132a. 模式變種var. laxicymosum

32. 花梗被短柔毛;萼距長20 mm

132b. 毛序疏傘翠雀花var. pilostachyum

25. 花序總狀。

33. 萼距直，稍向下彎曲，或成直角向下彎曲。

34. 萼片比距長;花梗長5～9 cm

108. 長梗翠雀花D. longipedicellatum

34. 萼片與距近等長或較短。

35. 萼片與距近等長。

36. 萼距向斜上方伸展，長10～13 mm

110. 直距翠雀花D. orthocentrum

36. 萼距不向上方伸展。

37. 莖被短柔毛;萼距水平方向伸展，長12～16 mm。

38. 花梗有少數(shù)黃色腺毛;萼片長12～18 mm，距長10～16 mm;子房被短柔毛

111. 小瓣翠雀花D. micropetalum

38. 花梗無黃色腺毛;萼片長12 mm，距長9～12 mm;心皮無毛

109. 擬直距翠雀花D. orthocentroides

37. 莖被長硬毛;萼距稍向下彎曲，長20 mm

112. 毛莖翠雀花D. hirticaule

39. 花梗無毛

112a. 模式變種var. hirticaule

39. 花梗有黃色腺毛

112b. 腺毛毛莖翠雀花var. mollipes

35. 萼片比距短。

40. 小苞片與花鄰接或近鄰接，并貼伏于花上。

41. 小苞片狹條形

122. 陰地翠雀花D. umbrosum

42. 小苞片寬0.5～1.1 mm。

43. 莖高達(dá)110 cm，被短糙伏毛;小苞片寬0.5～0.6 mm

122a. 模式變種var. umbrosum

43. 莖高達(dá)70 cm，被開展硬毛;小苞片寬0.6～1.1 mm

122b. 展毛陰地翠雀花var. hispidum

42. 小苞片寬1.5～1.8 mm

122c. 寬苞陰地翠雀花 var. drepanocentrum

41. 小苞片條狀披針形。

44. 莖被短糙伏毛;萼片外面被短柔毛

120. 滇川翠雀花 D. delavayi

45. 花梗被白色短糙伏毛和黃色短腺毛

120a. 模式變種var. delavayi

45. 花梗只被白色短糙伏毛

120b. 保山翠雀花var. baoshanense

44. 莖密被開展糙硬毛;萼片外面疏被糙硬毛

121. 須花翠雀花D. pogonanthum

46. 雄蕊無毛

121a. 模式變種var. pogonanthum

46. 雄蕊花絲每側(cè)有1～2根毛

121b. 毛蕊須花翠雀花var. lasiandrum

40. 小苞片多少與花分離，不貼伏花上。

47. 葉中央裂片不明顯3淺裂或不分裂。

48. 葉中央裂片菱形，邊緣中部之上具牙齒，中部之下全緣。

49. 萼片長9 mm，距長17 mm;莖被糙伏毛或無毛

113. 淡紫翠雀花D. handelianum

49. 萼片長15～16 mm，距長20～21 mm;莖無毛

114. 河南翠雀花 D. honanense

50. 花梗被白色短柔毛和黃色短腺毛

114a. 模式變種var. honanense

50. 花梗無黃色腺毛

114b. 毛梗河南翠雀花 var. piliferum

48. 葉中央裂片披針形，自基部之上至頂端有密集牙齒

115. 毛葉翠雀花D. hirtifolium

47. 葉中央裂片菱形，近中部明顯3淺裂，稀3深裂。

51. 葉中央裂片寬菱形，3深裂近中脈

129. 假深藍(lán)翠雀花D. pseudocyananthum

51. 葉中央裂片菱形，3淺裂。

52. 莖無毛。

53. 花梗無毛

136. 黑水翠雀花D. potaninii

54. 心皮無毛;蓇莢直，長圓形，長1.4～1.7 cm。

55. 小苞片寬0.2～0.3 mm

136a. 模式變種var. potaninii

55. 小苞片寬1～1.2 mm

136b. 寬苞黑水翠雀花var. latibracteolatum

54. 心皮有疏毛;蓇莢新月形，長0.5～1 cm

136c. 九峰山翠雀花var. jiufengshanense

53. 花梗有毛。

56. 花梗密被黃色短腺毛

57. 總狀花序復(fù)雜;心皮無毛。

58. 草本高1.5～2 m;花梗被白色短糙伏毛和黃色短腺毛;萼片長11 mm，距長18 mm;退化雄蕊瓣片2裂超過其中部

125. 多花翠雀花D. polyanthum

58. 草本高約80 cm;花梗只被黃色短腺毛;萼片長7～10 mm，距長11～12 mm;退化雄蕊瓣片頂端微凹

126. 左貢翠雀花D. zuogongense

57. 總狀花序簡單;子房被柔毛

127. 川西翠雀花D. tongolense

59. 葉柄無毛

127a. 模式變種var. tongolense

59. 葉柄被長硬毛

127b. 毛柄川西翠雀花var. hirsutipes

56. 花梗只被白色短柔毛，無黃色腺毛;總狀花序復(fù)雜;草本高約1 m

128. 高莖翠雀花D. altissimum

52. 莖有毛。

60. 莖和花梗被白色短柔毛和黃色短腺毛

123. 米林翠雀花 D. sherriffii

60. 莖和花梗無黃色腺毛。

61. 莖被開展糙硬毛;退化雄蕊瓣片被黃色髯毛。

62. 總狀花序簡單;萼片外面被白色短柔毛和黃色短腺毛;退化雄蕊的爪細(xì)，比瓣狹約5倍

116. 峨眉翠雀花 D. omeiense

63. 萼片長12～16mm，距長20～26 mm

116a. 模式變種 var. omeiense

63. 萼片長4～8mm，距長7～18mm

116b. 小花峨眉翠雀花 var. micranthum

62. 總狀花序復(fù)雜;萼片外面只疏被黃色腺毛，無白色毛，長10 mm，距長20 mm;退化雄蕊的爪條形，比瓣片稍狹

119. 寬爪翠雀花 D. platyonychium

61. 莖被柔毛。

64. 莖被開展柔毛;總狀花序復(fù)雜;萼距長23 mm

124. 鹽源翠雀花 D. yanyuanense

64. 莖被反曲短柔毛;總狀花序簡單;萼距長17～18 mm。

65. 葉片3深裂至距基部2～4 mm處; 上萼片狹卵形， 13 × 6 mm; 退化雄蕊瓣片被黃色髯毛

117. 柔毛翠雀花 D. pubescens

65. 葉片3深裂至距基部6～7 mm處; 上萼片寬卵形，10 × 7 mm; 退化雄蕊瓣片被白色髯毛

118. 周城翠雀花 D. zhouchengense

33. 萼距向下成U字形或螺旋狀彎曲。

66. 莖無毛。

67. 花梗無毛

136. 黑水翠雀花 D. potaninii

67. 花梗有毛。

68. 花梗只被黃色短腺毛;葉片寬達(dá)16 cm，3深裂稍超過中部;小苞片條形

141. 汶川翠雀花 D. wenchuanense

68. 花梗被白色短柔毛和黃色短腺毛;葉片寬達(dá)12 cm，3深裂至近基部;小苞片絲形。

69. 小苞片長4.5～7 mm;萼距長12～21 mm

140. 擬川西翠雀花 D. pseudotongolense

69. 小苞片長1.5～4 mm;萼距長18～24 mm

138. 擬螺距翠花 D. bulleyanum

70. 子房疏被柔毛

138a. 模式變種 var. bulleyanum

70. 心皮無毛

138b. 光果擬螺距翠雀花 var. leiogynum

66. 莖有毛。

71. 心皮無毛

139. 貢嘎翠雀花 D. gonggaense

71. 子房有毛。

72. 莖被開展硬毛;葉片寬達(dá)10（～14） cm;萼片長15～18 mm

142. 螺距翠雀花 D. spirocentrum

72. 莖被開展短柔毛;葉片寬達(dá)5.4 cm;萼片長12～13 mm

143. 狹菱形翠雀花 D. angustirhombicum

22. 葉片3全裂。

73. 葉全裂片細(xì)裂，末回裂片條狀披針形

157. 塔城翠雀花 D. aemulans

73. 葉全裂片不細(xì)裂，末回裂片卵形或三角形。

74. 萼距長15 mm以下。

75. 莖、花梗和心皮均無毛

144. 疏花翠雀花 D. sparsiflorum

75. 莖和花梗有毛。

76. 草本高約100 cm;總狀花序復(fù)雜;萼距長8～9 mm

145. 狹序翠雀花 D. angustipaniculatum

76. 草本高達(dá)60～80 cm;總狀花序簡單;萼距長12～15 mm。

77. 葉中央裂片菱形，在中部3淺裂，或近羽狀分裂;子房被柔毛

146. 錯(cuò)那翠雀花 D. conaense

77. 葉中央裂片寬菱形，在中部之下3深裂;心皮無毛

147. 秀麗翠雀花 D. wangii

74. 萼距長20～28 mm。

78. 莖無毛。

79. 萼片無角狀突起。

80. 花梗無毛。

81. 草本高達(dá)110 （～150） cm;葉片寬達(dá)20 cm;總狀花序復(fù)雜;小苞片長2.5～3.5 mm;萼距直或向下彎曲

151. 秦嶺翠雀花 D. giraldii

81. 草本高達(dá)70 cm;葉片寬達(dá)8 cm;總狀花序簡單;小苞片長4～5 mm;萼距螺旋狀彎曲

152. 光莖翠雀花 D. glabricaule

80. 花梗有毛。

82. 花梗被白色短柔毛和黃色短腺毛;總狀花序簡單

148. 木里翠雀花 D. muliense

83. 小苞片披針形或披針狀條形，8～13 × 1.2～2 mm

148a. 模式變種 var. muliense

83. 小苞片近絲形，3.5～4 × 0.2 mm

148b. 小苞木里翠雀花 var. minutibracteolatum

82. 花梗只被白色短柔毛，無黃色腺毛;總狀花序復(fù)雜

153. 松潘翠雀花 D. sutchuenense

79. 萼片外回頂端之下具角狀突起。

84. 花梗被白色短柔毛; 萼片角狀突起極小， 不明顯， 距水平伸展或稍向下彎曲; 心皮3

153. 松潘翠雀花 D. sutchuenense

84. 花梗被黃色短腺毛;萼片角狀突起明顯，長2.5～3.5，距呈U字形或螺旋狀彎曲;心皮3（～4）

154. 彎距翠雀花 D. campylocentrum

78. 莖有毛。

85. 花梗無毛。

86. 葉片下面無毛;萼片無毛，距長28 mm

156. 大藏翠雀花 D. dazangense

86. 葉片下面有毛;萼片有毛，距長20 mm。

87. 莖下部被黃色硬毛;萼片內(nèi)面無毛

149. 壤塘翠雀花 D. rangtangense

87. 莖下部被白色硬毛;萼片內(nèi)面有疏毛

150b. 光序擬彎距翠雀花 D. pseudocampylocentrum var. glabripes

85. 花梗有毛。

88. 草本高50 cm;花梗疏被黃色短腺毛;小苞片與花分開;萼距長17～22 mm;子房有疏毛

150a. 擬彎距翠雀花 D. pseudocampylocentrum var. pseudocampylocentrum

88. 草本高50～90 （～100） cm;花梗密被白色短柔毛;小苞片與花鄰接;萼距長22～25 mm;心皮無毛

155. 三小葉翠雀花 D. trifoliolatum

5. 基生葉發(fā)育，稀不存在，或存在時(shí)在花期枯萎。

89. 葉片3淺裂或3深裂。

90. 種子近四面體形，沿縱棱具翅。

91. 萼片外面頂端之下具角狀突起。

92. 葉片3裂稍超過中部，末回裂片卵形或?qū)捜?，稀狹披針或披針狀條形。

93. 葉一回中央裂片3次分裂

166. 大理翠雀花 D. taliense

94. 葉末回裂片卵形或三角形。

95. 萼距基部粗2～3 mm。

96. 萼片外面被短柔毛。

97. 莖無毛。

98. 心皮無毛

166a. 模式變種var. taliense

98. 子房有毛

166b. 毛蕊大理翠雀花 var. glabrum

97. 莖下部密被短柔毛

166c. 毛莖大理翠雀花 var. pubicaule

96. 萼片外面被長硬毛

166d. 硬毛大理翠雀花 var. hirsutum

95. 萼距基部粗4～5 mm

166e. 粗距大理翠雀花 var. platycentrum

94. 葉末回裂片狹披針或披針狀條形;萼距比萼片長2倍

166f. 長距大理翠雀花 var. dolichocentrum

93. 葉的1回中央裂片1～2次分裂

167. 角萼翠雀花 D. ceratophorum

99. 萼片角狀突起長1～2 mm。

100. 葉片寬達(dá)10 cm，末回裂片卵形或?qū)捜切巍?/p>

101. 萼片外面疏被短糙伏毛（毛長約0.4 mm），角狀突起長1.5～2 mm

167a. 模式變種 var. ceratophoram

101. 萼片外面被長硬毛（毛長約2 mm），角狀突起長1～1.5 mm

167c. 毛角萼翠雀花 var. hirsutum

100. 葉片寬8～19 cm，末回裂片條狀披針形;萼片角狀突起長1.5～2 mm

167d. 粗壯角萼翠雀花 var. robustum

99. 萼片角狀突起長在1 mm以下

167b. 短角萼翠雀花 var. brevicorniculatum

92. 葉片3深裂至近葉片基部處。

102. 花梗上部被黃色短腺毛和白色硬毛;萼片長10～12 mm;退化雄蕊瓣片被白色髯毛

169. 短萼翠雀花 D. brevisepalum

102. 花梗無毛;萼片長14～22 mm;退化雄蕊瓣片被黃色髯毛。

103. 莖無毛;萼片長18～22 mm，角狀突起長1 mm，距長24 mm

168. 擬角萼翠雀花 D. ceratophoroides

103. 莖下部被糙伏毛; 萼片長14～16 mm， 角狀突起長1～1.5 mm， 距長27～28 mm

170. 擬長距翠雀花 D. dolichocentroides

91. 萼片無角狀突起。

104. 葉片3裂達(dá)或稍超過中部。

105. 花梗無毛

162. 鉤距翠雀花 D. hamatum

105. 花梗被白色短柔毛和黃色短腺毛。

106. 莖近頂部被黃色短腺毛;萼片長9 mm，距長14～17 mm;心皮無毛

163. 寬菱形翠雀花 D. latirhombicum

106. 莖無毛;萼片長15～19 mm，距長20～22 mm;子房被柔毛

165. 寧朗山翠雀花 D. ninglangshanicum

104. 葉片3深裂至近基部處。

107. 萼片長13～19 mm。

108. 花梗無黃色腺毛。

109. 花梗上部具2小苞片。

110. 總狀花序長2～6.5（～16）cm;子房密被柔毛。

111. 總狀花序長16 cm;花梗長3.5～10 cm

174. 巖瓦翠雀花 D. yanwaense

111. 總狀花序長2～6.5 cm;花梗長1.4～3.2 cm。

112. 萼距長25～27 mm;退化雄蕊瓣片2半裂，被白色髯毛

172. 維西翠雀花 D. weixiense

112. 萼距長18 mm;退化雄蕊瓣片頂端微凹，被黃色髯毛

186. 洱源翠雀花 D. lankongense

110. 總狀花序長10～26 mm;子房只腹縫線有毛，其他部分無毛

187. 密距翠雀花 D. pycnocentrum

109. 花梗在頂端具2條形小苞片

164. 擬陰地翠雀花 D. umbrosoides

108. 花梗被黃色短腺毛和白色短柔毛;萼距長16～18 mm。

113. 莖無毛;雄蕊花絲有毛。

113′. 總狀花序長22 cm;退化雄蕊瓣片倒心形，寬7 mm;花藥無毛

182. 倒心片翠雀花 D. obcordatilimbum

113′. 總狀花序長12～15 cm;退化雄蕊瓣片長圓形，寬3 mm;花藥背面有毛

179. 毛藥翠雀花D. lasiantherum

113. 莖有毛; 花絲無毛。

113″. 莖被短柔毛，有1葉，基生葉約7;退化雄蕊瓣片2裂稍超過中部，被白色髯毛;子房被短柔毛

171. 缺刻翠雀花D. incisolobulatum

113″. 莖被糙伏毛，有數(shù)葉，基生葉在花期枯萎;退化雄蕊瓣片不分裂或2微裂，被黃色髯毛;心皮無毛

173. 甘孜翠雀花D. kantzeense

107. 萼片長20～30 mm。

114. 葉柄鞘發(fā)育良好，寬8～15 mm

180. 鞘柄翠雀花 D. coleopodum

114. 葉柄鞘很狹窄或不存在。

115. 花梗無毛。

116. 莖被短柔毛;萼片長9～14 mm，距長17～24 mm;退化雄蕊瓣片2半裂

178. 云南翠雀花 D. yunnanense

116. 莖無毛;萼片長18 mm，距長26～30 mm;退化雄蕊瓣片不分裂

185. 涼山翠雀花 D. liangshanense

115. 花梗有毛。

117. 花梗有黃色腺毛。

118. 莖被硬毛;葉片兩面疏被糙伏毛;萼距長17～20 mm

175. 永寧翠雀花 D. yongningense

118. 莖被短柔毛;葉片無毛;萼距長22 mm

176. 紙葉翠雀花 D. pergameneum

117. 花梗無黃色腺毛。

119. 草本高3～8 cm;花梗被白色短柔毛;萼片長15～16 mm，距長25 mm

181. 先骕翠雀花 D. hui

120. 葉片的中裂片具3齒;莖頂端具1～2花

181a. 模式變種var. hui

120. 葉片的中裂片具3～5齒;莖頂端具1～5（～10）花

181b. 多裂葉翠雀花var. lobulibundum

119. 草本高11～18 cm;花梗被黃色短柔毛;萼距長28～29 mm。

121. 莖下部疏被開展短柔毛;小苞片長5 mm;退化雄蕊瓣片頂端圓形，不分裂，全緣

183. 寧蒗翠雀花 D. pseudohamatum

121. 莖被反曲短柔毛;小苞片長（5～）7.5～9 mm;退化雄蕊瓣片頂端微凹或2微裂

184. 玉龍山翠雀花 D. yulongshanicum

90. 種子具密集橫向排列的小鱗片; 葉片3裂稍超過中部; 萼片長13～15 mm， 距長20～22 mm

188. 會(huì)澤翠雀花 D. hueizeense

89. 葉片3全裂。

122. 種子近四面體形，沿縱棱具狹翅。

123. 花序總狀。

124. 上萼片船形。

125. 葉片基部深心形，末回裂片三角形或狹披針形;萼距比萼片短，筒狀鉆形，長11～18 mm

190. 麗江翠雀花 D. likiangense

125. 葉片基部近截形，末回裂片條形;萼距比萼片長，近筒形，長21～27（～30）mm

191. 川甘翠雀花 D. souliei

124. 上萼片多少展開，不呈船形。

126. 葉的末回裂片卵形，三角形或狹披針形。

127. 萼片比距長或與距近等長。

128. 莖和花梗被短柔毛;萼片比距長，長17～23 mm，距長12～16 mm

192. 墨脫翠雀花 D. medogense

128. 莖和花梗無毛;萼片和距均長10～12 m

193. 德浚翠雀花 D. yuanum

127. 萼片比距短。

129. 莖密被開展短柔毛

210. 威海翠雀花 D. weihaiense

129. 莖被貼伏短柔毛或硬毛，或無毛。

130. 花梗被黃色短腺毛和白色短柔毛

204b. 銳裂翠雀花 D. thibeticum var. laceratilobum

130. 花梗無黃色腺毛。

131. 花梗無毛;葉均基生

199. 短葶翠雀花 D. breviscaposum

131. 花梗有毛。

132. 莖和心皮無毛;萼距長15 mm

194. 絲苞翠雀花 D. filibracteolatum

132. 莖和心皮有毛。

133. 退化雄蕊瓣片不分裂。

134. 萼距長24～28 mm;退化雄蕊瓣片頂端嚙蝕狀

201. 囊謙翠雀花 D. nangchienense

134. 萼距長16～24 mm;退化雄蕊瓣片頂端微凹。

135. 小苞片狹長圓形，長11～18 mm;萼距長達(dá)19 mm

195. 威寧翠雀花 D. weiningense

135. 小苞片狹條形或鉆形，長2.5～6 mm;萼距長達(dá)24 mm

200. 金沙翠雀花 D. majus

133. 退化雄蕊瓣片2裂。

136. 葉均基生， 中央全裂片近楔形， 1回分裂， 近頂端2淺裂或具3牙齒

198. 察隅翠雀花 D. chayuense

136. 葉基生并莖生，中央全裂片2回分裂。

137. 莖被反曲短柔毛;葉中央全裂片楔狀菱形;小苞片長圓形，長9～12 mm;萼片長15～18 mm，距長14～20 mm

196. 太白翠雀花 D. taipaicum

137. 莖被開展硬毛; 葉中央全裂片菱形; 小苞片條形， 長7～8 mm; 萼片長10～14 mm， 距長20～24 mm

197. 擬瀾滄翠雀花 D. pseudothibeticum

126. 葉的全裂片細(xì)裂，末回裂片披針狀條形、條形或狹條形。

138. 心皮5。

139. 花序軸和花梗被反曲短柔毛， 無黃色腺毛; 萼片長9 mm， 距長12 mm; 退化雄蕊瓣片頂端微凹; 心皮無毛

202. 五果翠雀花 D. sinopentagynum

139. 花序軸和花梗有黃色腺毛; 萼片長15 mm， 距長20～24 mm; 退化雄蕊瓣片2微裂; 子房被柔毛

203. 條裂翠雀花 D. pseudomosoynense

140. 花序軸和花梗被白色短柔毛和黃色短腺毛;上萼片外面頂端之下有1小角狀突起，距長20 mm;花瓣頂端被緣毛

203a. 模式變種 var. pseudomosoynense

140. 花序軸和花梗只被黃色短腺毛， 無白色短柔毛; 上萼片無角狀突起， 距長21～24 mm; 花瓣無毛

203b. 疏毛條裂翠雀花 var. subglabrum

138. 心皮3。

141. 退化雄蕊瓣片長圓形成倒卵狀長圓形。

142. 花序軸和花梗無毛。

143. 葉末回裂片寬6 mm; 總狀花序長25～45 cm; 萼距長13～15 mm; 退化雄蕊瓣片長4.6 mm， 2半裂

177. 擬云南翠雀花 D. pseudoyunnanense

143. 葉末回裂片寬1.6～3 mm;總狀花序長10～20 cm;萼距長25～28 mm;退化雄蕊長8 mm，2微裂

205. 光軸翠雀花 D. leiostachyum

142. 花序軸和花梗有毛。

144. 花序軸和花梗只被白色短柔毛;小苞片與花分開

208. 孟獲營翠雀花 D. mosoynense

144. 花序軸和花梗被白色短柔毛和黃色短腺毛;小苞片與花鄰接

204a. 瀾滄翠雀花 D. thibeticum var. thibeticum

141. 退化雄蕊瓣片寬倒卵形或近圓形。

145. 莖和花梗被短柔毛和短腺毛。

146. 莖和花梗密被開展白色短柔毛和白色短腺毛

211. 煙臺(tái)翠雀花 D. chefoense

146. 莖和花梗密被貼伏白色短腺毛和黃色開展短腺毛。

147. 子房被短柔毛

209d. 腺毛翠雀花 D. grandiflorum var. glandulosum

147. 心皮無毛

209e. 光果翠雀花 D. grandiflorum var. leiocarpum

145. 莖和花梗只被柔毛，無腺毛。

148. 花序軸和花梗近無毛或有極稀疏毛。

149. 莖無毛或下部有稀疏硬毛;萼片與距近等長

207. 秋翠雀花D. autumnale

149. 莖被極稀疏開展短柔毛

206. 光序翠雀花 D. kamaonense

150. 萼距比萼片稍短，長12～16 mm

206a. 模式變種var. kamaonense

150. 萼距比萼片長，長18～25 mm

206b. 展毛光序翠雀花 var. glabrescens

148. 花序軸和花梗密被柔毛

209. 翠雀 D. grandiflorum

151. 莖和花梗只密被貼伏短柔毛。

152. 子房密被短柔毛

209a. 模式變種var. grandiflorum

152. 心皮無毛

209c. 安澤翠雀花 var. deinocarpum

151. 莖和花梗密被貼伏短柔毛，此外還有開展長柔毛

209b. 疏毛翠雀花 var. pilosum

123. 花序傘房狀，或花單朵頂生。

153. 心皮5。

154. 萼片比距短，下萼片不呈船形，不具角狀突起;退化雄蕊瓣片被黃色髯毛。

155. 萼距近筒形。

156. 莖只被白色短柔毛，無黃色腺毛

212. 寬距翠雀花 D. beesianum

157. 葉的中央全裂片和側(cè)全裂片的2回裂片均有小裂片。

158. 葉的末回裂片寬2～3.5 mm

212a. 模式變種var. beesianum

158. 葉的末回裂片寬3～7 mm

212b. 粗裂寬距翠雀花var. latisectum

157. 葉的中央全裂片和側(cè)全裂片的2回裂片均無小裂片

212c. 輻裂寬距翠雀花var. radiatifolium

156. 莖被白色短柔毛和黃色短腺毛

213d. 粗距藍(lán)翠雀花D. caeruleum var. crassicalcaratum

155. 萼距鉆形。

159. 葉的末回裂片寬1.5～2.5（～4） mm;萼距基部粗3 mm;退化雄蕊瓣片寬倒卵形或近圓形，頂端全緣或微凹

213. 藍(lán)翠雀花 D. caeruleum

160. 莖和葉柄被貼伏短柔毛。

161. 葉片寬達(dá)5 cm

213a. 模式變種 var. caeruleum

161. 葉片寬達(dá)12 cm

213c. 大葉藍(lán)翠雀花 var. majus

160. 莖和葉柄被開展短柔毛

213b. 鈍裂藍(lán)翠雀花 var. obtusilobum

159. 葉的末回裂片寬0.5～1.5 mm; 萼距基部粗2 mm; 退化雄蕊片斜卵形， 2半裂

215. 細(xì)距翠雀花 D. lagarocentrum

154. 萼片與距近等長; 下萼片近頂端呈船形， 有時(shí)具小角狀突起; 退化雄蕊瓣片被白色髯毛

214. 永登翠雀花 D. yongdengense

153. 心皮3。

162. 花梗被白色短柔毛和黃色短腺毛。

163. 花梗全部被白色短柔毛和黃色短腺毛;退化雄蕊瓣片寬倒卵形。

164. 草本高30～80 cm; 莖被貼伏短柔毛; 傘房花序有3～12花; 萼距長20～25 mm

217. 康定翠雀花 D. tatsienense

165. 葉末回裂片寬1.5～2.5（～4） mm;萼片內(nèi)面無毛

217a. 模式變種 var. tatsienense

165. 葉末回裂片寬0.5～1.5 mm;萼片內(nèi)面有短毛

217b. 班瑪翠雀花 var. chinghaiense

164. 草本高10～16.5 cm; 莖被開展短柔毛; 傘房花序有（1～）2（～3）花; 萼距長33～38 mm

218. 矮翠雀花 D. pumilum

163. 花梗只在頂端被白色短柔毛和黃色短腺毛， 其他部分無毛; 退化雄蕊瓣片寬長圓形， 長13 mm

219. 碩片翠雀花 D. grandilimbum

162. 花梗無黃色腺毛。

166. 退化雄蕊的爪狹楔形，比瓣片短，比瓣片窄2倍

220. 波密翠雀花 D. pomeense

166. 退化雄蕊的爪細(xì)，與萼片近等長，比瓣片窄5～6倍。

167. 葉基生并莖生。

168. 葉片分裂程度小， 末回裂片三角形或狹卵形; 種子橢圓體形， 其縱棱不明顯， 無翅

216. 谷地翠雀花 D. davidii

168. 葉片強(qiáng)度分裂，全裂片細(xì)裂，末回裂片披針狀條形、條形或狹條形。

169. 草本高90～100 cm;葉末回裂片狹條形，寬1.5～1.8 mm;傘房花序有2～5花

221. 擬藍(lán)翠雀花 D. pseudocaeruleum

169. 草本高達(dá)22～40 cm。

170. 莖和花梗被開展短柔毛; 葉的末回裂片寬3.5 mm; 萼距長25 mm; 傘房花序有2 花

222. 軟毛翠雀花 D. mollipilum

170. 莖和花梗被貼伏短柔毛。

171. 草本高30～40 cm;花組成具2～3花的傘房花序。

172. 葉片寬達(dá)9.5 cm， 側(cè)全裂片3回細(xì)裂， 末回條形裂片寬2.5～4 mm; 退化雄蕊瓣片頂端微凹

209f. 房山翠雀花 D. grandiflorum var. fangshanense

172. 葉片寬達(dá)5 cm，側(cè)全裂片1回深裂，末回條形裂片寬1～2 mm;退化雄蕊瓣片頂端2微裂

225. 狹裂翠雀花 D. lagarolobum

171. 草本高12～22 cm;花單朵頂生;退化雄蕊瓣片2半裂

226. 李恒翠雀花 D. lihengianum

167. 葉全部基生。

173. 花葶被短柔毛， 頂端有1（～2）花; 萼片長16 mm， 距長22～24 mm; 退化雄蕊瓣片2淺裂， 被白色髯毛

223. 花葶翠雀花 D. sinoscaposum

173. 花葶被硬毛， 頂端有1花; 萼片長23～24 mm， 距長27 mm; 退化雄蕊瓣片2深裂稍超過中部， 被黃色髯毛

224. 巖生翠雀花 D. saxatile

122. 種子具橫向的條紋或小鱗片;萼距長達(dá)25 mm。

174. 種子近四面體形， 具橫向的條紋; 葉片3深裂至近基部處; 總狀或近傘房狀花序， 具少數(shù)花， 軸和花梗無毛

227. 長距翠雀花 D. tenii

174. 種子倒卵球形， 具橫向排列的小鱗片。

175. 草本高約100 cm; 莖無毛;萼距長15～22 mm; 花序近傘房狀， 有2～3花， 花梗無毛; 心皮無毛

228. 水城翠雀花 D. shuichengense

175. 草本高約60 cm; 莖被貼伏短柔毛; 花序傘房狀， 有2花， 花梗被貼伏短柔毛; 子房密被短柔毛

229. 多枝翠雀花 D. maximowiczii

Key to species and varieties

1. Petals and staminodes yellowish or yellowish-white; sepals blue or purple.

2. Leaf blades 3-sect; flowers solitary and terminal; sepals longer than spur; petals yellowish-white; staminode limbs yellowish-white， undivided; carpels 5

159. Delphinium huangzhongense

2. Leaf blades 3-parted; flowers in racemes; staminode limbs 2-lobed; carpels 3 or 1.

3. Sepals nearly as long as spur; petals and staminodes yellowish; carpels 3.

4. Flowers smaller， in compound racemes; sepals 6-7 mm long， with apexes by right angle incurved; carpels glabrous

160. D. zhengyianum

4. Flowers larger， in simple racemes; sepals 14 mm long， not incurved; ovaries puberulous

161. D. neowentsaii

3. Sepals shorter than spur; petals and staminodes yellowish-white; carpel l， glabrous

189. D. monogynum

1. Petals and staminodes concolour with sepals， i. e. blue and purple， rarely white.

5. Basal leaves absent， or present， then withered by anthesis.

6. Seed with 3 longitudinal edges and along edges narrow-winged.

7. Carpels 4-6.

8. Leaf blades 3-sect; carpels 5

98. D. kingianum

9. Stems hairy; leaf segments at apex acute.

10. Stens densely white-puberulous and yellow-glandular-puberulous.

11. Ovaries densely puberulous

98a. var. kingianum

11. Ovaries only on ventral sutures hairy， elsewhere glabrous

98b. var. leiocarpum

10. Stems densely white-puberulous and mixed with a few yellow glandular hairs

98c. var. eglandulosum

9. Stems glabrous; leaf segments at apex long acuminate

98d. var. acuminatissimum

8. Leaf blades 3-parted; carpels 4-6

99. D. humilius

7. Carpels 3.

12. Leaf blades 3-sect; sepals longer than spur

100. D. caudatolobum

12. Leaf blades 3-parted; sepals nearly as long as spur or shorter than it.

13. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous

103. D. gyalanum

13. Raceme rachis and pedicels laching yellow glandular hairs.

14. Stems and pedicels glabrous.

15. Herbs up to 140 cm tall; sepals 14-15 mm long; staminode limbs 2-lobed

106. D. cheilanthum

16. Pedicels retrorsely white-puberulous

106a. var. cheilanthum

16. Pedicels spreadingly yellow-puberulous

106b. var. pubescens

15. Herbs up to 20-30 cm tall; sepals 15-20 mm long.

17. Leaf primary lateral lobes 2-cleft; sepals 15-20 mm long， spur 18-24 mm long; staminode limbs suborbicular， 2-lobed

104. D. latilimbum

17. Leaf primary lateral lobes 2-parted exceeding the middle of lobe; sepals 15-18 mm long， spur 20 mm long; staminode limbs broad-elliptic， at apex emarginate

107. D. yuchuanii

14. Stems and pedicels hairy， rarely stems subglabrous （D. hsinganense）.

18. Sepals on both surfaces hairy， nearly as long as spur or slightly longer than it

101. D. pachycentrum

19. Sepals oblong

101a. var. pachycentrum

19. Sepals lanceolate

101b. var. lancisepalum

18. Sepals inside glabrous.

20. Compound racemes; bracteoles appressed on flower， narrow-lanceolate， 6-11 mm long; sepals 13 mm long; staminode limbs obovate-oblong， white-barbate

102. D. pachycentroides

20. Simple racemes; bracteoles remote from flower， subulate， 3-5 mm long; sepals 14-17 mm long; staminode limbs orbicular-ovate， yellow-barbate

105. D. hsinganense

6. Seeds densely transversely squamulose or striate.

21. Leaf blades broad-cordate， undivided or inconspicuously 3-lobulate

158. D. calthifolium

21. Leaf blades pentagonal or reniform， 3-lobed， 3-parted or 3-sect.

22. Leaf blades 3-lobed or 3-parted.

23. Leaf blades 3-lobed not exceeding the middle of blade.

24. Leaf blades 3-lobed not to the middle of blade， lobes not imbricate among themselves; inflorescences racemose， 10 cm long; pedicels white-puberulous and yellow-glandular-puberulous

135. D. sinovitifolium

24. Leaf blades 3-cleft to the middle of blade， with lobes imbricate among themselves; inflorescences corymbose， 6 cm long; pedicels glabrous

134. D. maoxianense

23. Leaf blades 3-parted exceeding the middle of blades.

25. Inflorescences corymbose.

26. Stems glabrous.

27. Sepals abaxially below apex corniculate

133. D. furcatocornutum

27. Sepals not corniculate.

28. Sepal spur spirally curved

137. D. bonvalotii

28. Sepal spur slightly recurved

130. D. eriostylum

26. Stems hairy.

29. Stems puberulous; leaf blades up to 6 cm broad

131. D. pingwuense

29. Stems hispid; leaf blades up to 12 cm broad.

30. Pedicels with yellow glandular hairs

130. D. eriostylum

31. Pedicels only yellow-glandular-puberulous; ovaries sparsely pubescent

130a. var. eriostylum

31. Pedicels white-puberulous and yellow-glandular-puberulous; carpels glabrous

130b. var. hispidum

30. Pedicels lacking yellow-glandular hairs

132. D. laxicymosum

32. Pedicels glabrous; sepal spur 23-27 mm long

132a. var. laxicymosum

32. Pedicels puberulous; sepal spur 20 mm long

132b. var. pilostachyum

25. Inflorescences racemose.

33. Sepal spur straight， slightly or by right angle recurved.

34. Sepals longer than spur; pedicels 5-9 cm long

108. D. longipedicellatum

34. Sepals nearly as long as or shorter than spur.

35. Sepals nearly as long as spur.

36. Sepal spur upwards spreading， 10-13 mm long

110. D. orthocentrum

36. Sepal spur not upwards spreading.

37. Stems puberulous; sepal spur horizontally spreading， 12-16 mm long.

38. Pedicels with few yellow glandular hairs; sepals 12-18 mm long， spur 10-16 mm long; ovaries puberulous

111. D. micropetalum

38. Pedicels lacking yellow glandular hairs; sepals 12 mm long，spur 9-12 mm long; carpels glabrous

109. D. orthocentroides

37. Stems hispid; sepal spur slightly recurved， 20 mm long

112. D. hirticaule

39. Pedicels glabrous

112a. var. hirticaule

39. Pedicels with yellow glandular hairs

112b. var. mollipes

35. Sepals shorter than spur.

40. Bracteoles contiguous or nearly contiguous to the flower and appressed to it.

41. Bracteoles narrow-linear

122. D. umbrosum

42. Bractaoles 0.5-1.1 mm broad.

43. Stems up to 110 cm tall， strigillose; bracteoles 0.5-0.6 mm broad

122a. var. umbrosum

43. Stems up to 70 cm tall， hispid; bracteoles 0.6-1.1 mm broad

122b. var. hispidum

42. Bracteoles 1.5-1.8 mm broad

122c. var. drepanocentrum

41. Bracteoles linear-lanceolate.

44. Stems strigillose; sepals outside puberulous

120. D. delavayi ? ? ? ? ? ? 45. Pedicels white-strigillose and yellow-glandular-puberulous

120a. var. delavayi

45. Pedicels only white-strigllose

120b. var. baoshanense

44. Stems hispid; sepals outside sparsely hispid

121. D. pogonanthum

46. Stamens glabrous

121a. var. pogonanthum

46. Stamen filaments at each side on margin with 1-2 hairs

121b. var. lasiandrum

40. Bracteoles remote from flower and not appressed to it.

47. Leaf central bobe inconspicuously 3-lobed or undivided.

48. Leaf central lobe rhombic， on margin above the middle densely dentate and below the middle entire.

49. Sepals 9 mm long， spur 17 mm long; stems strigose or glabrous

113. D. handelianum

49. Sepals 15-16 mm long， spur 20-21 mm long; stems glabrous

114. D. honanense

50. Pedicels white-puberulous and yellow-glandular-puberulous

114a. var. honanense

50. Pedicels only white-puberulous

114b. var. piliferum

48. Leaf central lobe lanceolate， on margin from above base to apex densely dentate

115. D. hirtifolium

47. Leaf central lobe rhombic， near the middle conspicuously 3-lobed， rarely 3-parted.

51. Leaf central lobe wide rhombic， near midrib 3-parted

129. D. pseudocyananthum

51. Leaf central lobe rhombic， 3-lobed.

52. Stems glabrous.

53. Pedicels glabrous

136. D. potaninii

54. Carpels glabrous; follicles straight， oblong， 1.4-1.7 cm long.

55. Bracteoles 0.2-0.3 mm broad

136a. var. potaninii

55. Bracteoles 1-1.2 mm broad

136b. var. latibracteolatum

54. Carpels with sparse hairs; follicles lunated， 0.5-1 cm long

136c. var. jiufengshanense

53. Pedicels hairy.

56. Pedicels densely yellow-glandular-puberulous.

57. Racemes compound; carpels glabrous.

58. Herbs 1.5-2 m tall; pedicels white-strigillose and yellow-glandular-puberulous; sepals 11 mm long， spur 18 mm long; staminode limbs 2-parted exceeding the middle of limb

125. D. polyanthum

58. Herbs ca. 80 cm tall; pedicels only yellow-glandular-puberulous; sepals 7-10 mm long， spur 11-13 mm long; staminode limbs at apex emarginate

126. D. zuogongense

57. Racemes simple; ovaries pubescent

127. D. tongolense

59. Petioles glabrous

127a. var. tongolense

59. Petioles hirsute

127b. var. hirsutipes

56. Pedicels only white-puberulous， lacking yellow glandular hairs; racemes compound; herbs ca. 1 m tall

128. D. altissimum

52. Stems hairy.

60. Stems and pedicels white-puberulous and yellow-glandular-puberulous

123. D. sherriffii

60. Stems and pedicels lacking yellow glandular hairs.

61. Stems hispid; staminode limbs yellow-barbate.

62. Racemes simple; sepals outside white-puberulous and yellow-glandular-puberulous; staminode claws slender， ca. 5 times narrower than limbs

116. D. omeiense

63. Sepals 12-16 mm long， spur 20-26 mm long

116a. var. omeiense

63. Sepals 4-8 mm long， spur 7-18 mm long

116b. var. micranthum

62. Racemes compound; sepals outside only with yellow glandular hairs， 10 mm long， spur 20 mm long; staminode claws linear， slightly narrower than limbs

119. D. platyonychium

61. Stems pubescent.

64. Stems spreading-pubescent; racemes compound; sepal spur 23 mm long

124. D. yanyuanense

64. Stems retrorsely puberulous; racemes simple; sepal spur 17-18 mm long.

65. Leaf blades 3-parted to 2-4 mm above base; upper sepal narrow-ovate， 13 × 6 mm; staminode limbs yellow-barbate

117. D. pubescens

65. Leaf blades 3-parted to 6-7 mm above base; upper sepal broud-ovate， 10 × 7 mm; staminode limbs white-barbate

118. D. zhouchengense

33. Sepal spur by U-form or spirally recurved.

66. Stems glabrous.

67. Pedicels glabrous

136. D. potaninii

67. Pedicels hairy.

68. Pedicels only yellow-glandular-puberulous; leaf blades up to 16 cm broad， 3-parted slightly exceeding the middle of blade; bracteoles linear

141. D. wenchuanense

68. Pedicels white-puberulous and yellow-glandular-puberulous; leaf blades up to 12 cm broad， 3-parted to near the base of blade; bracteoles filiform.

69. Bracteoles 4.5-7 mm long; sepal spur 12-21 mm long

140. D. pseudotongolense

69. Bracteoles 1.5-4 mm long; sepal spur 18-24 mm long

138. D. bulleyanum

70. Ovaries spasely pubescent

138a. var. bulleyanum

70. Carpels glabrous

138b. var. leiogynum

66. Stems hairy.

71. Carpels glabrous

139. D. gonggaense

71. Ovaries hairy.

72. Stems hispid; leaf blades up to 10（-14） cm broad; sepals 15-18 mm long

142. D. spirocentrum

72. Stems spreading-puberulous; leaf blade up to 5.4 cm broad; sepals 12-13 mm long

143. D. angustirhombicum

22. Leaf blades 3-sect.

73. Leaf segments dissected， with ultimate lobes linear-lanceolate

157. D. aemulans

73. Leaf segments not dissected， with ultimate lobes ovate or triangular.

74. Sepal spur less than 15 mm long.

75. Stems，pedicels and carpels glabrous

144. D. sparsiflorum

75. Stems and pedicels hairy.

76. Herbs ca. 100 cm tall; racemes compound; sepal spur 8-9 mm long

145. D. angustipaniculatum

76. Herbs 60-80 cm tall; racemes simple; sepal spur 12-15 mm long.

77. Leaf central lobe rhombic， at the middle 3-lobed or subpinnately divided; ovaries pubescent

146. D. conaense

77. Leaf central broad-rhombic， below the middle 3-parted; carpels glabrous

147. D. wangii

74. Sepal spur 20-28 mm long.

78. Stems glabrous.

79. Sepals not corniculate.

80. Pedicels glabrous.

81. Herbs up to 110 （-150） cm tall; leaf blades up to 20 cm broad; racemes compound; bracteoles 2.5-3.5 mm long; sepal spur straight or recurved

151. D. giraldii ? ? ? ? ?81. Herbs up to 70 cm tall; leaf blades up to 8 cm broad; racemes simple; bracteoles 4-5 mm long; sepal spur spirally recurved

152. D. glabricaule

80. Pedicels hairy.

82. Pedicels white-puberulous and yellow-glandular-puberulous; racemes simple

148. D. muliense

83. Bracteoles lanceolate or lanceolate-linear， 8-13 × 1.2-2 mm

148a. var. muliense

83. Bracteoles subfiliform， 3.5-4 × 0.2 mm

148b. var. minutibracteolatum

82. Pedicels only white-puberulous; racemes compound

153. D. sutchuenense

79. Sepals outside below apex corniculate.

84. Pedicels white-puberulous; sepal horn-like projections minute， inconspicuous; spur horizontally spreading or slightly recurved

153. D. sutchuenense

84. Pedicels yellow-glandular-puberulous; sepals with conspicuous horn-like projections 2.5-3.5 mm long， spur by U-form or spirally recurved

154. D. campylocentrum

78. Stems hairy.

85. Pedicels glabrous.

86. Leaf blades abaxially glabrous; sepals glabrous， spur 28 mm long

156. D. dazangense

86. Leaf blades abaxially hairy; sepals hairy， spur 20 mm long.

87. Stems below yellow-hispid; sepals inside glabrous

149. D. rangtangense

87. Stems below white-hispid; sepals inside with sparse hairs

150b. D. pseudocampylocentrum var. glabripes

85. Pedicels hairy.

88. Herbs ca. 50 cm tall; pedicels sparsely yellow-glandular-puberulous; bracteoles remote from flower; sepal spur 17-22 mm long; staminode limbs yellow-barbate; ovaries with sparse hairs

150a. D. pseudocampylocentrum var. pseudocampylocentrum

88. Herbs 50-90 （-100） cm tall; pedicels densely white-puberulous; bracteoles contiguous to flower; sepal spur 22-25 mm long; staminode limbs not barbate; carpels glabrous

155. D. trifoliolatum

5. Basal leaves well developed， rarely absent or present but withered by anthesis.

89. Leaf blades 3-lobed or 3-parted.

90. Seeds subtetrahedral， along edges narrow-winged.

91. Sepals outside below apex corniculate.

92. Leaf blades 3-parted slightly exceeding the middle of blade，with ultimate lobes ovate or broad-triangular， rarely narrow-lanceolate or lanceolate-linear.

93. Leaf central lobe thrice divided

166. D. taliense

94. Leaf ultimate lobes ovate or triangular.

95. Sepal spur at base 2-3 mm across.

96. Sepals outside puberulous.

97. Stem glabrous.

98. Carpels glabrous

166a. var. taliense

98. Ovaries puberulous

166b. var. glabrum

97. Stems below densely puberulous

166c.var. pubicaule

96. Sepals outside hirsute

166d. var. hirsutum

95. Sepal spur at base 4-5 mm across

166e. var. platycentrum

94. Leaf ultimate lobes narrow-lanceolate or lanceolate-linear; sepal spur 2 times longer than sepals

166f. var. dolichocentrum

93. Leaf central lobe once or twice divided

167. D. ceratophorum

99. Sepals with horn-like projections 1-2 mm long.

100. Leaf blades up to 10 cm broad， ultimate lobes ovate or triangular.

101. Sepal outside sparsely strigillose （hairs ca. 0.4 mm long）， with horn-like projections 1.5-2 mm long

167a. var. ceratophorum

101. Sepals outside hirsute （hairs ca. 2 mm long）， with horn-like projections 1-1.5 mm long

167c. var. hirsutum

100. Leaf blades 8-19 cm broad， ultimate lobes linear-lanceolate; sepal horn-like projections 1.5-2 mm long

167d. var. robustum

99. Sepal horn-like projections less than 1 mm long

167b. var. brevicorniculatum

92. Leaf blades 3-parted to near the base of blade.

102. Pedicels above yellow-glandular-puberulous and white-hispid; sepals 10-12 mm long; staminode limbs white-barbate

169. D. brevisepalum

102. Pedicels glabrous; sepals 14-22 mm long; staminode limbs yellow-barbate.

103. Stems glabrous; sepals 18-22 mm long， horn-like projections 1 mm long， spur 24 mm long

168. D. ceratophoroides

103. Stems below strigose; sepals 14-16 mm long， horn-like projections 1-1.5 mm long， spur 27-28 mm long

170. D. dolichocentroides

91. Sepals not corniculate.

104. Leaf blades 3-cleft or 3-parted slightly exceeding the middle of blade.

105. Pedicels glabrous

162. D. hamatum

105. Pedicels white-puberulous and yellow-glandular-puberulous.

106. Stems near apex yellow-glandular-puberulous; sepals 9 mm long， spur 14-17 mm long; carpels glabrous

163. D. latirhombicum

106. Stems glabrous; sepals 15-19 mm long; ovaries pubescent

165. D. ninglangshanicum

104. Leaf blades 3-parted to near the base of blade.

107. Sepals 13-19 mm long.

108. Pedicels lacking yellow glandular hairs.

109. Pedicels above 2-bracteolate.

110. Racemes 2-6.5（-16） cm long; ovaries densely pubescent.

111. Racemes 16 cm long; pedicels 3.5-10 cm long

174. D. yanwaense

111. Racemes 2-6.5 cm long; pedicels 1.4-3.2 cm long.

112. Sepal spur 25-27 mm long; staminode limbs 2-cleft， white-barbate

172. D. weixiense

112. Sepal spur 18 mm long; staminode limbs at apex emarginate， yellow-barbate

186. D. lankongense

110. Racemes 10-26 cm long; ovaries only on ventral sutures hairy， elsewher glabrous

187. D. pycnocentrum

109. Pedicels on apex 2-bracteolate （bracteoles linear）

164. D. umbrosoides

108. Pedicels yellow-glandular-puberulous and white-puberulous; sepal spur 16-18 mm long.

113. Stem glabrous; stamen filaments hairy.

113′. Racemes 22 cm long; staminode limbs obcordate， 7 mm broad; anthers glabrous

182. D. obcordatilimbum

113′. Racemes 12-15 cm long; staminode limbs oblong， 3 mm broad; anthers abaxially hair

179. D. lasiantherum

113. Stems hary; filaments glabrous.

113″. Stems puberulous， 1-leaved; basal leaves ca. 7; staminode limbs 2-parted slightly exceedling the middle of limb， white-barbate; ovaries puberulous

171. D. incisolobulatum

113″. Stems strigose， several-leaved; basal leaves withered by anthesis; staminode limbs undivided or at apex 2-lobulate， yellow-barbate; carpels glabrous

173. D. kantzeense

107. Sepals 20-30 mm long.

114. Leaf sheaths well developed， 8-15 mm broad

180. D. coleopodum

114. Leaf sheaths narrow or absent.

115. Pedicels glabrous.

116. Stems puberulous; sepals 9-14 mm long， spur 17-24 mm long; staminode limbs 2-cleft

178. D. yunnanense

116. Stems glabrous; sepals 18 mm long， spur 26-30 mm long; staminode limbs undivided

185. D. liangshanense

115. Pedicels hairy.

117. Pedicels with yellow glamdular hairs.

118. Stems hispid; leaf blades on both surfaces sparsely strigose; sepal spur 17-26 mm long

175. D. yongningense

118. Stems puberulous; leaf blades glabrous; sepal spur 22 mm long

176. D. pergameneum

117. Pedicels lacking yellow glandular hairs.

119. Herbs 3-8 cm tall; pedicels white-puberulous; sepals 15-16 mm long， spur 25 mm long

181. D. hui

120. Leaf blade with central lobe 3-dentate; stem apex 1-2-flowered

181a. var. hui

120. Leaf blade with central lobe 3-5-dentate; stem apex 1-5（-10）-flowered

181b. var. lobulibundum

119. Herbs 11-18 cm tall; pedicels yellow-puberulous; sepal spur 28-29 mm long.

121. Stems below sparsely spreading-puberulous; bracteoles 5 mm long; staminode limbs at apex rounded and entire

183. D. pseudohamatum

121. Stems retrorse-puberulous; bracteoles （5-）7.5-9 mm long; staminode limbs at apex emarginate or 2-lobulate

184. D. yulongshanicum

90. Seeds densely transversely squamulose; leaf blades 3-parted slightly exceeding the middle of blade; sepals 13-15 mm long， spur 20-22 mm long

188. D. hueizeense

89. Leaf blades 3-sect.

122. Seeds subtetrahedral， along longitudinal edges narrow-winged.

123. Inflorescences racemose.

124. Upper sepal eymbiform.

125. Leaf blades at base deep-cordate， ultimate lobes triangular or narrow-lanceolate; sepal spur shorter than sepals， tubular-subulate， 11-18 mm long

190. D. likiangense

125. Leaf blades at base subtruncate， ultimate lobes linear; sepal spur longer than sepals， subtubular， 21-27（-30） mm long

191. D. souliei

124. Upper sepal more or less flat， not cymbiform.

126. Leaf ultimate lobes ovate， triangular or narrow-lanceolate.

127. Sepals longer than spur or nearly as long as it.

128. Stems and pedicels puberulous; sepals longer than spur， 17-23 mm long; spur 12-16 mm long

192. D. medogense

128. Stems and pedicels glabrous; sepals and spur 10-12 mm long

193. D. yuanum

127. Sepals shorter than spur.

129. Stems densely spreading-puberulous

210. D. weihaiense

129. Stems appressed-puberulous， hispid or glabrous.

130. Pedicels yellow-glandular-puberulous and white-puberulous

204b. D. thibeticum var. laceratilobum

130. Pedicels lacking yellow glandular hairs.

131. Pedicels glabrous; leaves all basal

199. D. breviscaposum

131. Pedicels hairy.

132. Stems and carpels glabrous; sepal spur 15 mm long

194. D. filibracteolatum

132. Stems and carpels hairy.

133. Staminode limbs undivided.

134. Sepal spur 24-28 mm long; staminode limbs at apex erose

201. D. nangchienense

134. Sepal spur 16-24 mm long; staminode limbs at apex emarginate.

135. Bracteoles narrow-oblong， 11-18 mm long; sepal spur up to 19 mm long

195. D. weiningense

135. Bracteoles narrow-linear or subulate， 2.5-6 mm long; sepal spur up to 24 mm long

200. D. majus

133. Staminode limbs 2-lobed.

136. Leaves all basal， central segment subcuneate， once divided， near apex 3-lobed or 3-dentate

198. D. chayuense

136. Leaves basal and cauline， central segment thrice divided.

137. Stems retrorse-puberulous; leaf central segment cuneate-rhombic; bracteoles oblong， 9-12 mm long; sepals 15-18 mm long， spur 14-20 mm long

196. D. taipaicum

137. Stems spreading-hispid; leaf central segment rhombic; bracteoles linear， 7-8 mm long; sepals 10-14 mm long， spur 20-24 mm long

197. D. pseudothibeticum

126. Leaf segments dissected， with ultimate lobes lanceolate-linear， linear or narrow-linear.

138. Carpels 5.

139. Raceme rachis and pedicels retrorse-puberulous， lacking yellow glandular hairs; sepals 9 mm long， spur 12 mm long; staminode limbs at apex emarginate; carpels glabrous

202. D. sinopentagynum

139. Raceme rachis and pedicels with yellow glandular hairs; sepals 15 mm long， spur 20-24 mm long; staminode limbs at apex 2-lobulate; ovaries pubescent

203. D. pseudomosoynense

140. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous; upper sepal outside below apex with a small horn-like projection， spur 20 mm long; petals at apex ciliate

203a. var. pseudomosoynense

140. Raceme rachis and pedicels only yellow-glandular -puberulous， lacking white hairs; upper sepal lacking horn-like projection， spur 21-24 mm long; petals glabrous

203b. var. subglabrum

138. Carpels 3.

141. Staminode limbs oblong or obovate-oblong.

142. Raceme rachis and pedicels glabrous.

143. Leaf ultimate lobes 6 mm broad; racemes 25-45 cm long; sepal spur 13-15 mm long; staminode limbs 4.6 mm long， 2-cleft

177. D. pseudoyunnanense

143. Leaf ultimate lobes 1.6-3 mm broad; racemes 10-20 cm long; sepal spur 25-28 mm long; staminode limbs 8 mm long， 2-lobulate

205. D. leiostachyum

142. Raceme rachis and pedicels hairy.

144. Raceme rachis and pedicels only white-puberulous; bracteoles remote from flower

208. D. mosoynense

144. Raceme rachis and pedicels white-puberalous and yellow-glandular-puberulous; bracteoles contiguous to flower

204a. D. thibeticum var. thibeticum

141. Staminode limbs broad-obovate or suborbicular.

145. Stems and pedicels puberulous and glandular-puberulous.

146. Stems and pedicels densely spreading-white-puberulous and white-glandular- pube-rulous

211. D. chefoense

146. Stems and pedicels densely appressed-white-puberulous and spreading-yellow-puberulous.

147. Ovaries puberulous

209d. D. grandiflorum var. glandulosum

147. Carpels glabrous

209e. D. grandiflorum var. leiocarpum

145. Stems and pedicels only puberulous， not glandular-puberulous.

148. Raceme rachis and pedicels subglabrous or with very sparse hairs.

149. Stems glabrous or below sparsely hispid; sepals nearly as long as spur

207. D. autumnale

149. Stems with very sparse spreading hairs

206. D. kamaonense

150. Sepal spur slightly shorter than sepals， 12-16 mm long

206a. var. kamaonense

150. Sepal spur longer than sepals， 18-25 mm long

206b. var. glabrescens

148. Raceme rachis and pedicels with dense hairs

209. D. grandiflorum

151. Stems and pedicels only densely puberulous.

152. Ovaries densely appressed-puberulous

209a. var. grandiflorum

152. Carpels glabrous

209c. var. deinocarpum

151. Stems and pedicels densely appressed-puberulous， and with some spreading longer hairs

209b. var. pilosum

123. Inflorescences corymbose， or flowers solitary and terminal.

153. Carpels 5.

154. Sepals shorter than spur， lower sepal not， not corniculate; staminode limbs yellow-barbate.

155. Sepal spur subtubular.

156. Stems only white-puberulous， lacking yellow glandular hairs

212. D. beesianum

157. Leaf central segment and secondary lobes of lateral segments all with lobules.

158. Leaf ultimate lobes 2-3.5 mm broad

212a. var. beesianum

158. Leaf ultimate lobes 3-7 mm broad

212b. var. latisectum

157. Leaf central segment and secondary lobes of lateral segments all lacking lobules

212c. var. radiatifolium

156. Stems white-puberulous and yellow-glandular-puberulous

213d. D. caeruleum var. crassicalcaratum

155. Sepal spur subulate.

159. Leaf ultimate lobes 1.5-2.5（-4） mm broad; sepal spur at base 3 mm across; staminode limbs broad-obovate or suborbicular， at apex entire or emarginate

213. D. caeruleum

160. Stems and petioles appressed-puberulous.

161. Leaf blades up to 5 cm broad

213a. var. caeruleum

161. Leaf blades up to 12 cm broad

213c. var. majus

160. Stems and petioles spreading-puberulous

213b. var. obtusilobum

159. Leaf ultimate lobes 0.5-1.5 mm broad; sepal spur at base 2 mm across; staminode limbs white-barbate

215. D. lagarocentrum

154. Sepals nearly as long as spur; lower sepals near apex cymbiform， sometimes minutely 1-corniculate; staminode limbs white-barbate

214. D. yongdengense

153. Carpels 3.

162. Pedicels white-puberulous and yellow-glandular-puberulous.

163. Pedicels totally white-puberulous and yellow-glandular-puberulous; staminode limbs broad-obovate.

164. Herbs 30-80 cm tall; stems appressed-puberulous; corymbs 3-12-flowered; sepal spur 20-25 mm long

217. D. tatsienense

165. Leaf ultimate lobes 1.5-2.5（-4） mm broad; sepals inside glabrous

217a. var. tatsienense

165. Leaf ultimate lobes 0.5-1.5 mm broad; sepals inside with short hairs

217b. var. chinghaiense

164. Herbs 10-16.5 cm tall; stems spreading-puberulous; corymbs （1-）2（-3）-flowered; sepal spur 33-38 mm long

218. D. pumilum

163. Pedicels only on apex white-puberulous and yellow-glandular-puberulous， elsewhere glabrous; staminode limbs broad-oblong， 13 mm long

219. D. grandilimbum

162. Pedicels lacking yellow glandular hairs.

166. Staminode claws narrow-cuneate， shorter than limbs and two times narrower than limbs

220. D. pomeense

166. Staminode claws slender， nearly as long as limbs and 5-6 times narrower than them.

167. Leaves basal and cauline.

168. Leaf blades weakly divided， with ultimate lobes triangular or narrow ovate; seeds ellipsoidal， with inconspicuous longitudinal edges and not winged

216. D. davidii

168. Leaf blades strongly divided， with segments dissected and ultimate lobes lanceolate- linear， linear or narrow-linear.

169. Herbs 90-100 cm tall; leaf ultimate lobes narrow-linear， 1.5-1.8 mm broad; corymbs 2-5-flowered

221. D. pseudocaeruleum

169. Herbs up to 22-40 cm tall.

170. Stems and pedicels spreading-puberulous; leaf ultimate lobes 3.5 mm broad; sepal spur 25 mm long; corymbs 2-flowered

222. D. mollipilum

170. Stems and pedicels appressed-puberulous.

171. Herbs 30-40 cm tall; flowers in 2-3-flowered corymbs.

172. Leaf blades up to 9.5 cm broad， lateral segments thrice dissected， with ultimate lobes 2.5-4 mm broad; staminode limbs at apex emarginate

209f. D. grandiflorum var. fangshanense

172. Leaf blades up to 5 cm broad， lateral segments once parted， with ultimate lobes 1-2 mm broad; staminode limbs at apex 2-lobulated

225. D. lagarolobum

171. Herbs 12-22 cm tall; flowers solitary and terminal; staminode limbs 2-cleft

226. D. lihengianum

167. Leaves all basal.

173. Scapes puberulous， on apex 1（-2）-flowered; sepals 16 mm long， spur 22-24 mm long; staminode limbs 2-lobed

223. D. sinoscaposum

173. Scapes hispid， on apex 1-flowered; sepals 23-24 mm long， spur 27 mm long; staminode limbs 2-parted slightly exceeding the middle of limb

224. D. saxatile

122. Seeds transversely striate or squamulose.

174. Seed subtetrahedral， transversely striate; leaf blades 3-parted to near base of blade; racemes subcorymbose， few-flowered

227. D. tenii

174. Seeds obovoid， transversely squamulose.

175. Herbs ca. 100 cm tall; stems glabrous; inflorescences subcorymbose， 2-3-flowered， pedicels glabrous; carpels glabrous

228. D. shuichengense

175. Herbs ca. 60 cm tall， appressed-puberulous; inflorescences corymbose 2-flowered， pedicels appressed-puberuous; ovaries puberulous

229. D. maxinowiczii