王 勇
(陜西理工大學(xué)生物科學(xué)與工程學(xué)院,漢中 723000)
筆者在大巴山區(qū)考察時(shí),發(fā)現(xiàn)蘭科(Orchidaceae)山蘭屬(Oreorchis Lindley)植株正在開花。這些植株形態(tài)與山蘭(O.patens(Lindley)Lindley)十分相似,但在形態(tài)和物候上存在差異,可以區(qū)別。在營(yíng)養(yǎng)體形態(tài)方面,這些植株體型較大,假鱗莖(2~3 cm×1~2.5 cm)和葉片(30~40 cm×2~3.5 cm)均明顯大于山蘭(假鱗莖1~2 cm×0.5~1.5 cm,葉13~30 cm×0.4~2 cm);前者在假鱗莖頂端多數(shù)具3片葉,少數(shù)為2 片,而后者每個(gè)假鱗莖頂端通常只有1片葉,罕2片。此外,本次發(fā)現(xiàn)的開花植株葉柄頂端有明顯關(guān)節(jié),不同于《中國(guó)植物志》[1]、《Flora of China》[2],以及Pearce N.和Cribb P.[3]對(duì)山蘭葉的描述為“基部收狹為柄”。在花部形態(tài)方面,我們發(fā)現(xiàn)的開花植株的花朵唇瓣爪長(zhǎng)度約為總長(zhǎng)1/2,唇瓣側(cè)裂片直立,唇瓣中央縱褶片于末端(2唇瓣側(cè)裂片基部之間)膨大并匯合為船形附屬物,呈鮮艷的橙紅色,而山蘭花朵唇瓣爪較短,長(zhǎng)度通常僅為總長(zhǎng)約1/4,唇瓣側(cè)裂片向下鐮曲,唇瓣縱褶片脊?fàn)?,白色至淺黃色[1~6](見圖2),易于區(qū)別。同時(shí),在物候方面,本次發(fā)現(xiàn)的植株秋季生葉,經(jīng)冬不凋,翌年4月凋落,夏季無葉,5月開花,花期結(jié)束后假鱗莖即休眠,于9月在新生的假鱗莖頂端生出2~3片葉;不同于山蘭四季常綠,春季新葉長(zhǎng)出同時(shí)老葉凋落(周繇,私人通信)。根據(jù)以上特征,本次發(fā)現(xiàn)的山蘭屬植物是一個(gè)新種,特命名為巴山山蘭(Oreor?chis bashanensis),并描述如下。
Oreorchis bashanensis Wang Yong sp. nov.Fig.1:A-F
Species nova Oreorchis patens(Lindley)Lindley similis,a qua foilia 2-3,appendice carinatus aurantius supra medium inserta bene differt.
Plant terrestrial,up to 70 cm tall. Leaves fall when flowering,Pseudobulb ovoid to subellipsoid,2-3 cm×1-2.5 cm,2- or 3-noded,partly covered by scarious brown sheaths,persisting and forming a chain of pseudobulbs connected by a short rhizome.Leaf 3,or sometimes 2,linear or narrowly lanceolate,plicate,30-40 cm×2-3.5 cm,apex acuminate,gradually tapering at base. Petioles 5-8 cm long,sheathed. Inflorescence 35-70 cm,with 2 or 3 tubular sheaths below middle;rachis 10-20 cm,30-40 flowered;floral bracts narrowly lanceolate,3-5 mm.Flowers greenish brown,lip greenish brown on claw,midlobe and lateral lobs white with purple spots at apex;pedicel and ovary 10-15 mm;dorsal sepal narrowly obovate,ca. 9 mm×2 mm,apex slightly obtuse;lateral sepals narrowly ablong,ca. 8 mm×2 mm slightly falcate,apex acute;lip oblong-obovate,8 mm×3.5-5 mm,3-lobed below middle;long clawed at base(claw of midlobe ca. 4mm),lateral lobes erect,linear,ca.3 mm×0.7 mm,apex obtuse;apex of mid-lobe flabellate,up to 5.8 mm at widest point when spread,margins sinuate;disc with two longitudinal lamellae,up to 4 mm long,swell and fused into a carinate appendage,which is orange in color,projecting between the insertion points of lateral lobs.Column ca.4.5 mm,base thickened.Capsule oblong,ca.15 mm×7 mm.Fl.May-Jun,fr.Jul-Oct.
陜西(Shaanxi)寧強(qiáng)縣,紅石梁國(guó)家森林公園(Ning Qiang Xian,Hong Shi Liang National forest park),alt.1 800 m,落葉闊葉林下(under deciduous forest),2019-05-17 王勇(Wang Yong)20190517008(Holotype HZTC陜西理工大學(xué)植物標(biāo)本館)。
原記載世界山蘭屬共16 種,我國(guó)11 種[1~4,6],至此分別增為17 種和12 種,大巴山地區(qū)的蘭科植物的種數(shù)也由原記載的86種[7]增至87種。